Evanioidea Online

Flat in lateral view, with punctures usually conspicuous. Eye elliptical. Clypeus flat, protruding medially. Epistomal declivity of clypeus usually divergent, arched and short. Mandible with five teeth. Area surrounding antennal socket (torulus) slightly raised. Antenna elbowed, usually articulated above level of mesosoma, without some longer setae, inserted at level of midline of eye. Scape long (equal or greater than half eye height). Pedicel usually longer than wide. Flagellum with 11 meres. Flagellomeres each longer than wide, widening progressively (female) or evenly wide (male). First flagellomere shorter than remaining flagellomeres. Distalmost flagellomere longest. Subocular groove absent. Malar space at least 3 times greater than basal mandibular width. Gena wide, nitid. Occipital carina present as fine ridge and completed dorsally.

Higher than long, usually with inconspicuous punctures dorsally, slightly areolate ventrally. Mesosoma higher than head. Pronotum dorsally obscure in midline, with fine, irregular striae laterally. Mesoscutum raised in midline; lateral carina complete from anterior to posterior margin; notaulus conspicuously sinuous, posteriorly with dilation conspicuous, convergent and joined at scutellar groove; parapsidal furrow inconspicuous as a fine ridge, complete to anteroposteriorly region. Scutellar groove scrobiculate laterally. Metanotum scrobiculate, partially covered by scutellum anteriorly. Mesopleuron higher than wide, mostly without sculpture; anterodorsal corner scrobiculate; anteriorly nitid; posteriorly with an even scrobiculate line; anteroventral margin finely scrobiculate; ventrally usually mostly nitid. Metapleuron usually with inconspicuous sculpture, mostly areolate, usually polished ventrally. Propodeum short, less than half length of petiole, not raised to metanotum; dorsally without areolate to rugulose sculpturing laterally nitid; propodeal area ventral to petiole flat. Distance between mid and hind coxae less than distance between fore and mid coxae. Hind coxae mostly rugulose. Hind leg faintly imbricate, usually without some longer setae. Hind tibia and tarsus without conspicuous prominent spines. Internal tibial spur length almost two times greater than external tibial spur. Tarsal claw with apical greater than subapical tooth.

Fore wing with usually with six (Brazil sp. with seven) cells enclosed by tubular veins. 1st subdiscal cell usually open. First marginal cell digitiform. Ventral margin of apex arched, with angulation more opened. Abscissa between r-rs and 1R1 vein almost straight. 2R1 vein greater than first marginal cell length. R-m vein absent. Stigmal vein shorter, less than first submarginal cell length, narrowing posteriorly. 2Mb and 3M vein spectral. 3CU vein usually spectral. 1RS vein attached to Sc+R at stigmal vein. Hind wing without jugal lobe. Usually with three hamuli. M+CU vein absent.

Elliptical, or usually elliptical (depending on preservation). Petiole arched dorsally, usually without sculpture except for some punctures, without expansions laterally.

see species descriptions

The new genus-group name is a combination of a (Greek meaning, “absence of”), lobos (Greek meaning, “lobe”) and Evania (type genus of the family). The name is feminine.

Unknown.

Morphbank image collection (196254)
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n. gen. can be distinguished from Evaniella by the elliptical eye (closer to ovoid in Evaniella), body with less sculpture; propodeum laterally polished (areolate in Evaniella); fore wing with 6-7 enclosed cells (always seven in Evaniella); first marginal cell digitiform (subquadrangular in Evaniella), 2R1 vein length greater than first marginal cell (2R1 vein length less than first marginal cell in Evaniella); hind wing without jugal lobe (present in Evaniella and all other evaniid genera except some spp. in the Old World genus Prosevania), and metasoma elliptical (ovoid in Evaniella). The characters that most distinguish the species of this genus are: epistomal declivity of clypeus, petiole and metapleuron sculpture, notaulus configuration and number and shape cells of the fore wing.
This new genus, as with other genera in the family (see Deans and Huben (2003); Deans (2005); Deans (2006) and Kawada and Azevedo (2007)) is sexually dimorphic. Female eyes are smaller and elliptical (larger, softly bulging and more spherical eyes in male), antenna progressively enlarged (all segments the same diameter in male), metasoma with ovipositor protracted (genitalia concealed in male) and both sexes with elliptical metasoma. Some Evaniidae fossils, as Protoparevania Deans and Eovernevania Deans, share most of the synapomorphies that unite extant evaniids, except that they lack separated jugal lobes in the fore and hind wings (Figs. 1A, 3B). Possession of separated jugal lobes appears in only one Cretaceous (Turonian) evaniid genus, Newjersevania Basibuyuk, Quicke & Rasnitsyn (seen only in the hind wing of one specimen) (Basibuyuk et al. 2000). Lack of separated jugal lobes in n. gen. (and some Prosevania Kieffer) is considered to be a secondary loss.

Tropical rainforests of Central America (Costa Rica), south to Ecuador and Peru, east to Venezuela and southern Brazil.